Transylvanian Dinosaurs

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Transylvanian Dinosaurs Page 32

by David B Weishampel


  Zarski, M., Jakubowski, G., and Gawor-Biedowa, E. 1998. [The first Polish find of the Lower Paleocene crocodile Thoracosaurus Leidy, 1852: geological and palaeontological description.] Kwartalnik Geologiczny 42: 141–160. [In Polish.]

  Ziegler, A. M., Parrish, J. M., Yao J-.P., Gyllenhaal, E. D., Rowley, D. B., Totman Parrish, J., Nie, S.-Y., Bekker, A., and Hulver, M. L. 1993. Early Mesozoic phytogeography and climate. Philosophical Transactions of the Royal Society of London B, Biological Sciences 341: 297–305.

  Ziegler, A. M., Scotese, C. R., and Barrett, S. F. 1983. Mesozoic and Cenozoic paleogeographic maps. In: Brosche, P., and Sündermann, J. (eds.). Tidal Friction and the Earth’s Rotation, 2. Springer-Verlag, Berlin. Pp. 240–252.

  Ziegler, P. A. 1988. Evolution of the Arctic-North Atlantic and the Western Tethys. American Association of Petroleum Geologists Memoir 43. American Association of Petroleum Geologists, Tulsa, Oklahoma. 198 pp.

  Ziegler, P. A. 1990. Geological Atlas of Western and Central Europe. Shell Internationale Petroleum Maatschappij B.V., The Hague. 130 pp.

  Zweigel, P., Ratschbacher, L., and Frisch, W. 1998. Kinematics of an arcuate fold-thrust belt: the southern Eastern Carpathians (Romania). Tectonophysics 297: 177–207.

  INDEX

  Page numbers in italics indicate illustrations.

  Abel, Othenio, 10, 86, 87, 105

  Abelisaurus, 110

  acipenseriforms (chondrosteans), 82, 83

  actinistian fish, 154–56

  Actinopterygii, 82–83

  Acynodon, 67, 68, 70

  Adriatic Sea, 96

  Aegean Sea, 96

  Africa, 93, 95, 105, 112, 117, 169, 170, 179

  agamids, 73

  Ajkaceratops kozmai, 109

  Alamosaurus, 28, 169

  Albanerpeton, 81, 82

  Albanerpetontidae, 80, 81, 81

  Albania, 11–14

  Albertosaurus, 110

  Albian, 109, 217n15, 218n14

  Alectrosaurus, 110

  alligator lizards, 73

  Alligatoroidea, 71

  alligators, 19

  Allodaposuchus, 71, 179

  Allodaposuchus precedens, 67, 69, 69–71

  allopatry, 165

  Allosaurus, 32

  Alps, 94

  Alp-Tethys oceanic basin, 95

  Altirhinus, 52

  Alvarezsauridae, 31

  alvarezsaurids, 37

  ammonites, 98, 117

  Amniota, 60

  Ampelosaurus, 28, 107, 178, 179, 181

  Ampelosaurus atacis, 110, 180

  amphibians, 80–82, 107–9

  Amphisbaenia, 73

  Anabisetia, 45, 158, 160, 161, 169

  anatomy, 21–22, 114

  angiosperms, 102, 103, 188

  Anguimorpha, 72, 73

  Anhanguera, 64

  Ankylopollexia, 45

  Ankylosauria, 11, 25, 38–43, 141, 161

  Ankylosauridae, 39, 110

  Ankylosaurus, 39

  Anura, 80

  Anurognathus, 64

  Apatosaurus, 23, 133

  Apoda, 80

  Aptian, 95

  Apulia, 96–98, 105

  Apuseni Mountains, 15, 88, 94, 97, 101

  Arabia, 93

  Aralosaurus, 52

  Archaeopterydactyloidea, 64

  Archaeopteryx, 19, 30, 31

  Archosauria, 18, 19

  Argentina, 27, 117, 161

  Argentinosaurus, 117

  arthropods, 122

  Asia, 27

  comparisons with, 110, 169

  conditions in, 170, 171

  and Hadro-sauridae, 171, 174

  and migration, 182, 183, 189, 192

  and Transylvanian insularity, 112, 167

  Auca Mahuevo, 56–58

  Aude Valley, 174

  Australia, 169, 170

  Australopithecus afarensis, 18, 19

  Australopithecus africanus, 19

  Austria, 43, 109

  Austrochelyidae, 76

  Austrosaurus, 169

  Aves, 18, 33

  Avetheropoda, 31

  Avialae, 31, 37

  axolotl (Ambystoma mexicanum), 118, 119, 120

  Azhdarchidae, 64, 65, 67, 109, 141, 184

  Azhdarcho, 64

  Azhdarchoidea, 64

  Bactrosaurus, 52, 157, 171, 172

  Bactrosaurus johnsoni, 52

  Bagaceratops kozmai, 176

  Bakonydraco galaczi, 109

  Bakony Mountains, 176

  Balaur bondoc, 37, 38

  Balkan Mountains, 94

  Bărăbanţ, 15, 46, 73

  Barbatodon transsylvanicus, 77, 79, 81

  bat, 123

  Batrachia, 80

  Becklesius hoffstetteri, 72

  Becklesius nopcsai, 72

  beetle, dermestic, 104

  Belgium, 108

  Belgorod, 108, 109

  benettitaleans, 102

  Betasuchus bredai, 108

  Bicuspidon hatzegiensis, 72

  biogeography, 169–71, 184

  birds, 19, 30, 31, 36

  Black Sea, 1, 96, 108

  Boccaccio, Giovanni, 113

  body size, 117, 118

  and colonization, 142, 143, 185, 191, 192

  and croco-dilians, 118, 141

  and Euhadrosauria, 132

  and fauna, 86–87, 106

  and France, 137

  and Iguanodon, 45

  and Iguanodontia, 130

  and Mag-yarosaurus, 187

  and migration, 143–45, 187–89

  reasons for, 87, 118, 141, 142

  and selection, 185

  and Struthiosaurus, 187

  and Telmatosaurus, 128–32, 187, 189

  and Telmatosaurus transsylvanicus, 49

  and Thero-poda, 145

  and Zalmoxes, 46, 187. See also dwarfism; growth and development

  Bohemian Massif, 102

  Borod Basin, 32

  bowfin fish, 150

  Brachiosaurus, 28, 133

  Brachylophosaurus, 50

  Bradycneme, 36

  Broili, Ferdinand, 10

  Brookes, Richard, 114

  Buckland, William, 114–15, 116

  Buffetaut, Eric, 25, 65, 70, 175

  Bulgaria, 109

  Bunzel, Emmanuel, 39, 40, 61

  caecilians, 80

  Camarasaurus, 28

  Campanian, 95, 97

  Campanian–Maastrichtian interval, 95

  Camptosaurus, 25, 43–45, 129, 159, 160, 187

  Carcharodontosaurus, 30, 117

  Carnosauria, 31

  Carroll, Lewis, 194–95

  Casichelydia, 76

  Ćáslav, 109

  Caucasus, 93

  Caudata, 80

  Cenozoic era, 94, 153

  centipedes, 122

  Cephalopoda, 19

  Ceratopsia, 25, 111, 124, 161

  Ceratosauria, 31

  Cerna River, 87

  chameleons, 73

  character change, 131, 156–58

  rates of, 150, 151, 159–62, 171. See also evolution

  character optimization, 126–28, 130, 131, 170, 216n70

  Cherven Bryag, 109

  chimpanzee, 119, 120, 121

  Choceň, 60

  choristoderans, 108

  cladistics, 16, 18, 152, 168, 170, 171

  cladogenesis, 165

  cladogram, 18, 20, 21, 126–28, 151, 152

  clam, unionid, 104

  clubmoss (Lycopodiaceae), 103, 103

  Cochirleni, 204n4

  Codrea, Vlad, 15, 34, 77

  coelacanth, 148, 149, 154

  Coelophysoidea, 31

  Coelurosauria, 31, 37

  coevolution, 196, 197

  colonization, 162, 168, 186

  and body size, 142, 143, 185, 191–92

  and clutch size, 191–92

  and continental configurations, 182

  and diversity, 106

  in Europe, 185


  and founder effect, 166

  and loss of genetic variation, 186

  and sexual maturation, 186, 191

  success in, 167, 183, 184, 186, 192

  Comoros Islands, 148, 149, 162

  Compsognathidae, 31

  Compsognathus, 30, 31

  conifers, 102, 104

  continental drift, 89–91

  Cope, Edward Drinker, 117, 142

  Cope’s Rule, 117, 142

  Cornet, 204n4

  Courtenay-Latimer, Marjorie, 147–48, 154

  Cretaceous, 93

  ephemeral terrestrial habitats of, 159

  flora of, 102

  habitats of, 105

  and hadrosaurids, 52

  and isolation and colonization of Transylva-nian region, 145

  mammals in, 77

  and multituberculates, 77

  and ornitho-pods, 45

  and Pterosauria, 60, 64

  and sauropods, 28

  and Southern Carpathian Mountains, 94, 97

  and titano-saurs, 27

  Cretaceous-Tertiary boundary, 65, 77, 198

  Crimea, 108

  crocodilians, 3, 19, 98, 179, 180

  and Al-lodaposuchus precedens, 68–70

  body size of, 118, 141

  in Europe, 107–9

  eu-suchian, 109

  feeding by, 72

  growth of, 184

  social behavior of, 72

  Crocodilus affulvensis, 68

  Crocodylia, 18, 19, 71

  Crocodyloidea, 71

  Cryptodira, 76, 77

  Csiki, Zoltan, 27, 34, 44, 65, 124

  Ctenodactylidae, 64

  cuckoo (Cuculus canorus), 197

  Cuvier, Georges, 114

  cycads, 102

  Czechia, 60, 102, 109, 211n57

  Danube, 1

  Daspletosaurus, 110

  deer, 119

  Deinonychus, 30, 36

  Deinotherium giganteum, 114

  Densuş, 15

  Densuş-Ciula Formation, 37, 63, 67, 100–102, 159

  Devonian, 93

  Diacryptodira, 76

  Dietz, Robert S., 91, 92

  Dinornis maximus, 22

  Dinosauria, 18, 19, 22–25, 116, 142

  Diplodocoidea, 28, 110

  Diplodocus, 117, 133

  diversity, 16, 106, 109, 110, 165, 169

  Djungaripteroidea, 64

  dogs, 122

  Dollo, Louis, 10, 10

  Dollodon, 52

  donkeys, 119

  Doratodon, 67, 68, 70, 71

  dortokids, 77

  Drasković, Ludwig Graf, 11–12

  Dromaeosauridae, 31, 33, 34, 36, 36–38

  Dromaeosaurus, 35

  Dryomorpha, 45, 158

  Dryosaurus, 45, 159, 160, 187

  Dsungaripteridae, 64

  Dsungaripteroidea, 63, 64

  Dsungaripterus, 64

  dwarfism, 142–46, 184–87, 191, 192, 213n29

  and endocrine system, 87, 118, 141, 150

  and femoral structure, 124, 125

  and insular isolation, 86, 87, 142, 143. See also body size; growth and development

  Early Cretaceous, 169, 179

  migrations in, 183, 184

  and Neotethyan Ocean, 97

  and Severin Basin, 95

  and Telmatosaurus, 159

  Ebro High, 97

  Edmontonia, 39, 40

  eggs, 107, 108, 190–92

  angusticanalicu-late, 55

  and clutch size and colonization, 191, 192

  dendrospherulitic, 55

  and eggshell construction, 54, 55, 58

  hatchlings from, 55, 56

  nests of, 53–58

  elephants, 114, 119, 143–45

  Elopteryx, 36

  Elopteryx nopcsai, 32–33, 34, 36

  embryos, 55–58

  enantiornithine bird, 109

  England, 98, 115, 179, 211n57

  Eolambia, 52

  Eolambia caroljonesa, 171

  Euhadrosauria, 52, 131, 171–74, 189

  and body size, 132

  clutch size of, 192

  ghost lineage of, 157

  Euoplocephalus, 39

  euornithopod, 176

  Euramerica, 179, 180

  Euronychodon, 36, 36, 37

  Europe, 27, 93, 97, 174, 182

  and dinosaur distribution, 170, 171, 181

  and migration, 182, 185, 189

  as source area, 169, 178, 179

  Eusuchia, 70, 71, 109, 179

  evolution, 16–18, 21, 120, 142, 150, 151, 154, 156. See also character change; morphological change, rates of

  Fărcădin, 87

  Fennosarmatia, 97

  fern (Polypodiaceae), 103

  fern (pteridophytes), 102

  fern, filicalean (Gleicheniaceae), 103

  fern, filicalean (Polypodiaceae), 103

  Fontllonga hadrosaurid, 174

  foraminifera, 98

  France, 137, 169, 174, 175, 182

  and Ampelosaurus, 110, 178, 181

  fauna of, 106–10, 175

  Fritsch, Anton, 60

  frogs, 80, 81

  Gallodactylidae, 64

  gar (Lepisosteus oculatus), 83

  Gargoyleosaurus, 39

  Gasparinisaura, 45, 169

  Gastonia, 39

  Gavialoidea, 71

  Gavials, 19

  Gekkota, 73

  Germanodactylus, 64

  ghost lineage, 151–54, 171

  ghost lineage duration (GLD), 152, 153, 159, 160

  Giganotosaurus, 30, 32, 110, 117

  gigantism, 86, 87, 117, 141, 142, 145

  Gila monsters, 73

  ginkgoes, 102

  Gobisaurus, 39, 169

  Gondwana, 27, 93, 171, 178

  gorilla, 119, 120

  Gosau Beds (Austria), 43

  Gosau fauna, 61, 108

  Gould, Stephen J., 120, 146, 163, 199

  Grigorescu, Dan, 15, 33, 34, 65, 101, 124

  growth and development: acceleration of, 122, 123

  of crocodilians, 184

  of Hadrosauridae, 126

  of Hatzego-pteryx, 184

  among herbivores, 145

  of Hypsilophodon, 137–40

  lines of arrested (LAGs), 124

  of Magyaro-saurus, 126, 133–37, 141, 144, 146, 184

  of Magyarosaurus dacus, 125

  of Mammalia, 184

  of ornithopods, 125

  of Orodromeus, 137–40

  of pterosaurs, 184

  of Rhabdodon, 137–40

  of sauropods, 133–37

  and selection, 185

  of Struthiosaurus, 184

  of Tel-matosaurus, 126, 128–32, 135, 137, 141, 144, 146, 184

  of Telmatosaurus transsylvanicus, 124–25

  of Tenon-tosaurus, 138–40

  of Theropoda, 124, 184

  of titanosaurian sauropods, 126

  of titanosaurs, 137

  of turtles, 184

  of Tyrannosaurus rex, 124

  of Zalmoxes, 126, 137–40, 146

  of Zalmoxes ro-bustus, 125, 139–41, 184

  of Zalmoxes shqiperorum, 138–40, 184. See also body size; dwarfism

  Groza, Ioan, 15

  Gryposaurus, 53

  Gymnophiona, 80

  Hadrosauridae: cranial adornments of, 53

  dentition of, 45, 51, 129–32, 187–90, 192

  eggs of, 56

  in Europe, 108, 109, 174

  evolutionary relationships in, 52

  Fontllonga, 174

  growth of, 126

  and Ornithischia, 23

  and Para-rhabdodon, 174

  perinatal bones of, 58

  reproductive biology of, 190–92

  and Rhabdodontidae, 46

  and social behavior, 53

  source area of, 169, 171–72, 174

  in Spain, 176

  and Telmatosaurus, 132, 157

  and Telmatosau
rus transsylvanicus, 49–51

  and Tethyshadros insularis, 172

  Hadrosaurinae, 50–52, 131

  Hadrosauroidea, 52, 109

  Haeckel, Ernst, 120

  Haeckel’s Biogenic Law, 120

  Hainina, 77

  Harrison, Colin J. O., 33

  Haţeg: amphibians of, 81–82

  an-kylosaur material from, 40

  drom-aeosaurid from, 34, 35

  eggs from, 58

  fauna of, 15

  formations in, 99–101

  fossils from, 2, 25, 32, 36

  geography of, 98–105

  geology of, 100

  history of, 6

  and insular isolation, 84–88

  as island, 105–11

  mammals of, 77–81

  mesoeucrocodilians of, 67–73

  pal-eoecological context of, 16

  perinatal bones known from, 58

  and plate tectonics, 97

  pterosaurs from, 59–67

  Rhabdodon priscus from, 43

  river systems of latest Cretaceous in, 101

  squamates of, 72–77

  Struthiosaurus transylvanicus from, 38

  and Telmatosaurus transsylvanicus, 49

  turtles from, 73–77

  work in, 15

  Zalmoxes from, 46

  Hatzegobatrachus grigorescui, 81

  Hatzegopteryx, 64, 66, 72, 141, 184

  Hatzegopteryx thambema, 65–66

  Hennig, Willi, 18, 151

  Heptasteornis, 36

  Heptasteornis andrewsi, 33, 36

  Herrerasaurus, 24

  hesperornithids, 109

  heterochrony, 119–26, 134, 144, 146, 168

  hippos, dwarfed, 119

  historical contingency, 168, 190, 198, 199

  Holden, John C., 91, 92

  Hominidae, 18, 19

  Homo erectus, 19

  homology, 17–21

  homoplasies, 19–21

  Homo sapiens, 18, 19, 118

  Horner, Jack, 58, 190

  horsetails (sphenophytes), 102

  Huene, Friedrich von, 11, 11, 25, 27

  humans, 119–21

  Hunedoara County, 2

  Hungarosaurus tormai, 109, 176

  Hungary, 108, 109, 175, 176, 181

  Hylaeochampsa, 179

  Hylaeosaurus, 38, 169

  Hypacrosaurus, 190

  hypermorphosis, 122

  Hypsilophodon, 44, 45, 137–40, 169

  Hypsilophodon foxii, 139

  Hypsilophodontidae, 44

  Iberia, 97, 107, 182

  Iguania, 73

  iguanians, 73

  iguanids, 73

  Iguanodon: body size of, 45

  and hadrosaurids, 131, 187

  lower teeth of, 128, 129

  and migration, 169

  as or-nithischian, 23

  recognition of, 116

  and Rhabdodon, 43, 44

  and Rhabdo-dontidae, 46, 160

  teeth of, 130

  and Telmatosaurus, 49, 50

  Iguanodon bernissartensis, 52

  Iguanodontia: body size of, 130

  and Hadrosauridae, 171, 187

  and Or-nithopoda, 45

  rate of evolutionary change of, 160

  and Rhabdodontidae, 46, 159, 176

  teeth of, 130, 131

  and Telmatosaurus transsylvanicus, 157

 

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